INVERTEBRATE PALEONTOLOGY
FINAL EXAM 1999 Name__________________________
I. Identification: What term is defined or described by each of the following statements? (30)
1. Type of preservation by which soft tissues may be preserved as a thin film after volatiles (H & O) are driven off in a reducing environment. _________________________________
2. Ecological zone defined by the depth to which light penetrates the marine environment. ___________________________________
3. Larval form shared by several invertebrate phyla, including: annelids, arthropods, molluscs, brachiopods and bryozoans. ________________________________
4. Phosphatic, tooth-like microfossils found throughout the Paleozoic; now thought to be chordates. _________________________________________
5. Asexually-reproducing, sessile form of a coelenterate that undergoes alternation of generations. _________________________________
6. Term that describes a species that will tolerate only a narrow range of salinity conditions in the environment. _____________________________________
7. Lustrous shell substance composed of alternating layers of aragonite and organic material found in some mollusks; mother-of-pearl. __________________________
8. Habit of a graptolite rhabdosome characterized by repeated branching with secondary connections between branchs; thought to be primitive form for the group. ______________
9. "Head" of a crinoid from which the arms arise. ______________________
10. Root-like or massive structure that anchors crinoids (as well as some other sessile organisms) to the sea floor. ____________________________________
11. Term that describes organisms that lack nuclei or other organelles. _______________________________
12. Feeding structure that surrounds the mouth in bryozoans. __________________
13. Support structure attached to the lower valve in most brachiopods that attaches them to the substrate. ________________________________
14. Term for small calcareous or siliceous skeletal elements; used for many groups such as sponges or sea cucumbers. ________________________________
15. . Rasping feeding structure found in most mollusks. __________________________
II. Double entendre: The statements below describe a term that is used for structures that are similar in function but not homologous in different taxonomic groups of organisms. For each statement, name (a) the feature and (b) at least one other group that has a structure with the same name. (20)
1. Hinged structure that covers the aperture in cheilostome bryozoans.
a._________________________ b. ___________________________
2. Line that represents the external trace of the intersection of the septum with the outer wall of a cephalopod conch.
a. _________________________ b. ___________________________
3. The skeletal structure that houses an individual graptozoan zooid; arrayed along a stipe.
4. Disc-shaped, calcite plates that extend through the arms of ophiuroids.
5. Processes that occur on the pedicle valve of articulate brachiopods that key into sockets in the brachial valve.
a. ___________________________ b. ___________________________
III. Odd item out: From each list of 4 items, select the one that fits least well with the others. (10)
1. interval zone; assemblage zone; abundance zone; stereozone
2. phylogenetic; phenetic; dianetic; classical;
3. ascon; sycon; leucon; cephalon
4. stone canal; ring canal; radial canal; haversian canal
5. plankton; benthos; epifauna; infauna
IV. Functional morphology: For each of the following features tell which group is characterized by the feature and briefly describe its function. (20)
1. Aristotle's lantern
2. resilium
3. ascus
4. choanocytes (collar cells)
5. cardinal process
V. Each sentence below describes a wholly fossil group with no known living representatives. Name them. (10)
1. "Lacy" bryozoans with zoaria characterized by zooecia arranged on ribbons connected by dissepiments. ________________________________
2. Now thought to be sclerosponges, they consist of laminae separated by pillars. ________________________________
3. Squid-like cephalopods with a well-calcified, bullet-shaped, internalized shell. ________________________________
4. echinoderms that attach to the substrate by columns and that have canals through the plates of the theca that connect pore rhombs. _____________________________________
5. Organisms of unknown affinities that are constructed as nested, perforated cones. _____________________________
VI. Diagram and label the echinoderm water vascular (ambulacral) system. What is its function and expression in each of the major groups of echinoderms? (20)
VII. Answer 70 points worth of the following questions. Use diagrams or drawings as necessary or useful.
1. Explain the difference between rheophilic and rheophobic crinoids. (10)
2. Echinoderms can be identified as such even from small fragments in thin section. What is it about echinoderms that makes their skeletal remains so distinctive? (5)
3. As a field geologist given responsibility for mapping a certain area you encounter a formation bearing the following fossils. What is your best estimate of its age and how did you come to this conclusion? (5)
fusulinids
productid brachiopods
trepostome bryozoans
tabulate coral
trilobites
blastoids
4. From among the invertebrate groups we have studied, provide an example of groups that illustrate evolutionary convergence, and explain the reason for this phenomenon. (10)
5. What type of information is needed to employ graphic correlation in biostratigraphy according to the method developed by Shaw? How is it done? (10)
6. To what phylum are echinoderms most closely related? What features support this relationship? (10)
7. Explain how a fossil mold forms. (5)
8. Graptolites are widely considered to be chordates. What evidence from the graptolites themselves and from modern organisms supports this view? (5)
9. Jeffreys classifies one of the groups of homalozoan echinoderms as a chordate group he calls Calcichordata. What features of this echinoderm group might suggest such an interpretation? (5)
10. Reefs and reef-like structures have developed repeatedly throughout the history of life, but with different groups of organisms as the primary reef-builders. What groups have been the reef-builders and when throughout the Phanerozoic? (10)
11. What is the approximate stratigraphic range of each of the following groups?
a. trilobites b. ammonoids c. scleractinian corals d. fusulinids e. Inarticulata (15)
12. On a profile of the ocean basin from land to the deep ocean indicate the location of the different marine ecological zones distinguished by depth. (10)
13. Use a eurypterid to diagram the basic body plan of the chelicerates. (10)
14. As a field geologist given responsibility for mapping a certain area you encounter a formation bearing the following fossils. What is your best estimate of its age and how did you come to this conclusion? (5)
phacopid trilobites tabulate coral
strophomenid brachiopods ammonoids stromtoporoids
VIII. Select two of the following sets of taxa and construct a hypothesis of phylogenetic relationships among them in the form of a cladogram. Provide appropriate evidence for each grouping by indicating derived characters. (20)
Archaeocidaris
Diploporita
Asteroidea
Limpet
Crinoidea
Slug
clypeasteroid
Strophomena
sea urchin
Rugosa
Productidina
Blastoidea
Hydrozoa
Terebratulida
Neospirifer
Tetragraptus
blue-green algae
Archimedes
Composita
diatom
Pentamerus
chiton
Triticites
winkle
Syringopora
Queen Conch
crinoid
Lithostrotionella